A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species

Abstract

Hypselostomatidae is a large family of terrestrial pulmonate molluscs currently represented by 284 extant species, most confined to Southeast Asia. The current system of grouping species into genera is based on the morphology of the apertural barriers and the level of last whorl detachment. However, these characters overlap, challenging generic definitions. In this paper, these characters are evaluated and a novel classification proposed for hypselostomatid species belonging to the genera Anauchen (17 species), Bensonella (36 species), Boysidia (7 species, in part) and Hypselostoma (85 species). We assigned all species belonging to the genera Bensonella and Hypselostoma into two and four species groups respectively, which are characterised by combinations of morphological traits. Altogether 46 new species are described, seven species in Anauchen: A.crassus Gojšina, Hunyadi & Páll-Gergely, sp. nov., A.evanidus Gojšina & Páll-Gergely, sp. nov., A.grandiportus Gojšina, Grego & Páll-Gergely, sp. nov., A.obesus Gojšina, Hunyadi & Páll-Gergely, sp. nov., A.picasso Gojšina & Páll-Gergely, sp. nov., A.turritus Gojšina, Hunyadi & Páll-Gergely, sp. nov., A.jokaii Gojšina & Páll-Gergely, sp. nov.; 19 species in Bensonella: B.alycaeus Gojšina & Páll-Gergely, sp. nov., B.cardiostoma Gojšina, Vermeulen & Páll-Gergely, sp. nov., B.cristatissima Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.dha Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.dracula Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.exploda Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.fracta Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.microdentata Gojšina & Páll-Gergely, sp. nov., B.mitochondria Gojšina, Vermeulen & Páll-Gergely, sp. nov., B.mirabilis Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.montawa Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.multidentata Gojšina, A. Reischütz & Páll-Gergely, sp. nov., B.nitens Gojšina & Páll-Gergely, sp. nov., B.obex Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.perfecta Gojšina & Páll-Gergely, sp. nov., B.sericata Gojšina & Páll-Gergely, sp. nov., B.serrata Gojšina, Hunyadi & Páll-Gergely, sp. nov., B.spelaea Gojšina, Grego & Páll-Gergely, sp. nov., B.spinosa Gojšina, Hunyadi & Páll-Gergely, sp. nov.; 20 species in Hypselostoma: H.aquila Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.bubalus Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.circumcarinatum Gojšina, Auffenberg & Páll-Gergely, sp. nov., H.coriaceum Gojšina & Páll-Gergely, sp. nov., H.aenigma Gojšina, Grego & Páll-Gergely, sp. nov., H.fortunatum Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.fungus Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.geckophilum Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.iunior Gojšina & Páll-Gergely, sp. nov., H.ophis Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.platybasis Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.populare Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.sculpturatum Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.similare Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.sorormajor Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.sororminor Gojšina, Hunyadi & Páll-Gergely, sp. nov., H.torta Gojšina, Auffenberg & Páll-Gergely, sp. nov., H.vesovici Gojšina & Páll-Gergely, sp. nov., H.vicinum Gojšina, Auffenberg & Páll-Gergely, sp. nov., H.vujici Gojšina & Páll-Gergely, sp. nov. One replacement name is proposed: H.tertiusfrater Gojšina & Páll-Gergely, nom. nov. pro Boysidiasalpinx F. G. Thompson & Dance, 1983, non Hypselostomasalpinx (van Benthem Jutting, 1961) (originally described as Gyliotrachela). Gyliotrachela and Antroapiculus are both treated as junior synonyms of Hypselostoma. An additional 28 species and subspecies are reassigned to the synonymies of other taxa.